Robert G. Spiro, 1929 – 2015


Robert Gunter Spiro, professor emeritus at Harvard Medical School, died May 16, at age 86. His research focused on the kidney complications of diabetic patients.

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Alexander Rich, 1924 – 2015


Alexander Rich of the Massachusetts Institute of Technology biology department, a biophysicist who contributed pioneering research on the structure of DNA and RNA, died in April at age 90. Rich's research team discovered Z-DNA.

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David E. Ong, 1943 – 2015


David E. Ong of Vanderbilt University died in April at age 71. Ong's research on vitamin A changed the research community's understanding of the nutrient's significance.

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Nathan Aronson, 1940 – 2015


Nathan Aronson, former chairman of the University of South Alabama’s biochemistry department, died in March at the age of 74. His research primarily focused on diseases related to the digestion of tissue proteins.

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Irwin Rose, 1926 – 2015


Nobelist Irwin Rose, an assiduous enzymologist who helped to explain how cells destroy unwanted proteins, died in June. He was 88.

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